✓ Fits the Metroidvanias cluster, with adventure storytelling, Metroidvania exploration and precision platforming
≠ Its own twist: puzzle design, cozy, relaxing feel and fantasy world, while skipping roguelike runs
Owlboy is a pixel-art platformer game about an owl-humanoid boy called Otus. When his homeland was attacked by pirates, Otus starts an adventure to protect his home and reveal the secrets of his race. He makes new friends on the way. Each of them has special powers, useful during fights.
It predates Ultros by 8 years, with what that implies technically.